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Appropriate fractions were pooled, concentrated, and buffer exchanged into storage buffer (50 mM HEPES pH 7.5, 500 mM NaCl, 5% glycerol, 0.5 mM TCEP.) The MTRR construct without the FMN domain, residues 217-698 (MTRR ∆FMN ), was cloned from a codon-optimised sequence (Twist Bioscience) into pGTVL2 to produce a TEV-cleavable His-GST N-terminally tagged protein for bacterial expression